An Online Catalog of the Ants of the World
by Barry Bolton
Dorylinae
Leach, 1815
valid
Extant: 27 valid genera, 685 valid species, 118 valid subspecies
Fossil: 1 valid genus, 8 valid species
Dorylida Leach, 1815Leach, W. E. 1815. Entomology. Pp. 57-172 in: Brewster, D. (ed.) 1815. The Edinburgh encyclopedia. Edinburgh: William Blackwood, Volume 9. [1815-04] 126849: 147. Type-genus: Dorylus. AntWeb AntWiki HOL
- Dorylinae as family: Leach, 1815Leach, W. E. 1815. Entomology. Pp. 57-172 in: Brewster, D. (ed.) 1815. The Edinburgh encyclopedia. Edinburgh: William Blackwood, Volume 9. [1815-04] 126849: 147 [Dorylida (family-group name)]; Haliday, 1836Haliday, A. H. 1836. Descriptions, etc. of the Hymenoptera. Pp. 316-331 in: Curtis, J.; Haliday, A. H.; Walker, F. 1836. Descriptions, etc. of the insects collected by Captain P. P. King R. N., F. R. S., in the survey of the Straits of Magellan. Transactions of the Linnean Society of London 17:315-359. [1836-06-21] 125734: 331 [Dorylidae]; Swainson & Shuckard, 1840Swainson, W.; Shuckard, W. E. 1840. On the history and natural arrangement of insects. London: Longman, Brown, Green & Longman's, 406 pp. [1840-12] 129098: 175 [Dorylidae]; Shuckard, 1840aShuckard, W. E. 1840a. Monograph of the Dorylidae, a family of the Hymenoptera Heterogyna. Annals of Natural History 5:188-201. [1840-05] PDF 128626: 188 [Dorylidae]; Smith, 1859cSmith, F. 1859c. Catalogue of hymenopterous insects in the collection of the British Museum. Part VII. Dorylidae and Thynnidae. London: British Museum, 76 pp. [1859-07-23] PDF 128689: 1 [Dorylidae]; Mayr, 1866bMayr, G. 1866b. Diagnosen neuer und wenig gekannter Formiciden. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 16:885-908. [1866] PDF 127216: 895 [Dorylidae]; Smith, 1871bSmith, F. 1871b. A catalogue of the aculeate Hymenoptera and Ichneumonidae of India and the Eastern Archipelago. [concl.]. Journal of the Linnean Society of London. Zoology 11:349-415. [1871-12-20] PDF 128708: 225 [Dorylidae]; André, 1882aAndré, Ern. 1882a. Les fourmis. [part]. Pp. 49-80 in: André, Edm. 1882. 1881-1886. Species des Hyménoptères d'Europe et d'Algérie. Tome Deuxième. Beaune: Edmond André, 919 + 48 pp. [1882-01] 122251: 125 [Dorylidae]; Cresson, 1887Cresson, E. T. 1887. Synopsis of the families and genera of the Hymenoptera of America, north of Mexico, together with a catalogue of the described species, and bibliography. Transactions of the American Entomological Society, Suppl. Vol. 1887:1-351. [1887] PDF 123936: 93 [Dorylidae]; Emery, 1894hEmery, C. 1894h. [Untitled.]. Pp. 373-401 in: Ihering, H. 1894. von. Die Ameisen von Rio Grande do Sul. Berliner Entomologische Zeitschrift 39:321-447. [1894-10] 124581: 381 [Dorylidae]; Ashmead, 1905cAshmead, W. H. 1905c. A skeleton of a new arrangement of the families, subfamilies, tribes and genera of the ants, or the superfamily Formicoidea. Canadian Entomologist 37:381-384. [1905-11-01] PDF 122385: 381 [Dorylidae]; Ashmead, 1906Ashmead, W. H. 1906. Classification of the foraging and driver ants, or Family Dorylidae, with a description of the genus Ctenopyga Ashm. Proceedings of the Entomological Society of Washington 8:21-31. [1906-07-13] PDF 122380: 21 [Dorylidae]; Bernard, 1951cBernard, F. 1951c. Hyménoptères, super-famille des Formicoidea Ashmead 1905. Pp. 997-1104 in: Grassé, P. P. (ed.) 1951. Traité de Zoologie. Tome X. Fasc. II. Paris: Masson et Cie, pp. 976-1948. [1951] 122634: 1046 [Dorylidae]; Bernard, 1953bBernard, F. 1953b ("1952"). La réserve naturelle intégrale du Mt Nimba. XI. Hyménoptères Formicidae. Mémoires de l'Institut Français d'Afrique Noire 19:165-270. [1953-03] PDF 122637: 217 [Dorylidae].
- Dorylinae as subfamily of Dorylidae: Ashmead, 1905cAshmead, W. H. 1905c. A skeleton of a new arrangement of the families, subfamilies, tribes and genera of the ants, or the superfamily Formicoidea. Canadian Entomologist 37:381-384. [1905-11-01] PDF 122385: 381; Ashmead, 1906Ashmead, W. H. 1906. Classification of the foraging and driver ants, or Family Dorylidae, with a description of the genus Ctenopyga Ashm. Proceedings of the Entomological Society of Washington 8:21-31. [1906-07-13] PDF 122380: 25.
- Dorylinae as subfamily of Formicidae: Mayr, 1865Mayr, G. 1865. Formicidae. In: Novara Expedition 1865. Reise der Österreichischen Fregatte "Novara" um die Erde in den Jahren 1857, 1858, 1859. Zoologischer Theil. Bd. II. Abt. 1. Wien: K. Gerold's Sohn, 119 pp. [1865-12-14] PDF 127193: 16 [Dorylidae]; Emery, 1877bEmery, C. 1877b. Saggio di un ordinamento naturale dei Mirmicidei, e considerazioni sulla filogenesi delle formiche. Bullettino della Società Entomologica Italiana 9:67-83. [1877-03] PDF 124518: 70 [Dorylidae]; Forel, 1878cForel, A. 1878c. Études myrmécologiques en 1878 (première partie) avec l'anatomie du gésier des fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles 15:337-392. [1878-10] PDF 125003: 365 [Dorylidae]; Emery & Forel, 1879a: 465 [Dorylidae]; André, 1881cAndré, Ern. 1881c. Catalogue raisonné des Formicides provenant du voyage en Orient de M. Abeille de Perrin et description des espèces nouvelles. Annales de la Société Entomologique de France (6)1:53-78. [1881-07-22] PDF 122245: 64 [Dorylidae]; Forel, 1892kForel, A. 1892k. Les Formicides de l'Empire des Indes et de Ceylan. Part I. Journal of the Bombay Natural History Society 7:219-245. [1892-10-01] PDF 125041: 220 [Dorylidae]; Forel, 1893bForel, A. 1893b. Sur la classification de la famille des Formicides, avec remarques synonymiques. Annales de la Société Entomologique de Belgique 37:161-167. [1893-04-01>] PDF 125049: 163; Dalla Torre, 1893Dalla Torre, K. W. 1893. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). Leipzig: W. Engelmann, 289 pp. [1893] PDF 124002: 1; Forel, 1895bForel, A. 1895b. A fauna das formigas do Brazil. Boletim do Museu Paraense de Historia Natural e Ethnographia 1:89-139. [1895-04] PDF 125065: 118 [Dorylidae]; Emery, 1895lEmery, C. 1895l. Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8:685-778. [1895-10-18] PDF 124599: 764 [subfamily spelled Dorylini]; Emery, 1896eEmery, C. 1896e. Clef analytique des genres de la famille des Formicides, pour la détermination des neutres. Annales de la Société Entomologique de Belgique 40:172-189. [1896-05-02>] PDF 124609: 174; Forel, 1899BForel, A. 1899B. Formicidae. Dummy reference. Biologia Centrali-Americana Hym 3:1-169. 133007: 22; Forel, 1901aForel, A. 1901a. Les Formicides de l'Empire des Indes et de Ceylan. Part VIII. Journal of the Bombay Natural History Society 13:462-477. [1901-01-20] PDF 125105: 462; Forel, 1901dForel, A. 1901d. I. Fourmis mexicaines récoltées par M. le professeur W.-M. Wheeler. II. A propos de la classification des fourmis. Annales de la Société Entomologique de Belgique 45:123-141. [1901-04-30] PDF 125108: 139; Emery, 1901bEmery, C. 1901b. Notes sur les sous-familles des Dorylines et Ponérines (Famille des Formicides). Annales de la Société Entomologique de Belgique 45:32-54. [1901-03-07] PDF 124641: 36; Bingham, 1903Bingham, C. T. 1903. The fauna of British India, including Ceylon and Burma. Hymenoptera, Vol. II. Ants and Cuckoo-wasps. London: Taylor and Francis, 506 pp. [1903] PDF 122766: 1; Emery, 1910bEmery, C. 1910b. Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum 102:1-34. [1910-06-25] PDF 124696: 3; Wheeler, 1910aWheeler, W. M. 1910a. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp. [1910-03] 130044: 137; Arnold, 1915Arnold, G. 1915. A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae. Annals of the South African Museum 14:1-159. [1915-02-11] 122309: 110; Escherich, 1917Escherich, K. 1917. Die Ameise, Schilderung ihrer Lebensweise. Zweite verbesserte und vermehrte Auflage. Braunschweig: Friedrich Vieweg und Sohn, xvi + 348 pp. [1917] 124797: 2 [Dorylini]; Forel, 1917Forel, A. 1917. Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles 51:229-253. [1917-03-08] PDF 124997: 239; Bondroit, 1918Bondroit, J. 1918. Les fourmis de France et de Belgique. Annales de la Société Entomologique de France 87:1-174. [1918-12-24] PDF 122872: 14 [Dorylitae]; Wheeler, 1920Wheeler, W. M. 1920. The subfamilies of Formicidae, and other taxonomic notes. Psyche (Cambridge) 27:46-55. [1920-06] PDF 129953: 53; Wheeler, 1922Wheeler, W. M. 1922. Ants of the American Museum Congo expedition. New York: Bulletin of the American Museum of Natural History, 1139. 133015: 632; Clark, 1951Clark, J. 1951. The Formicidae of Australia. Vol. 1. Subfamily Myrmeciinae. Melbourne: CSIRO, 230 pp. [1951] PDF 123635: 16; Brown, 1954eBrown, W. L., Jr. 1954e. Remarks on the internal phylogeny and subfamily classification of the family Formicidae. Insectes Sociaux 1:21-31. [1954-03] PDF 123108: 28; Borgmeier, 1955Borgmeier, T. 1955. Die Wanderameisen der neotropischen Region. Studia Entomologica 3:1-720. [1955] PDF 122904: 51; all subsequent authors.
- Dorylinae as dorylomorph subfamily of Formicidae: Bolton, 2003Bolton, B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71:1-370. [2003-11-10] PDF 130789: 36, 147; Brady & Ward, 2005Brady, S. G.; Ward, P. S. 2005. Morphological phylogeny of army ants and other dorylomorphs (Hymenoptera: Formicidae). Systematic Entomology 30:593-618. [2005-10] PDF 130829: 593.
- Dorylinae as formicoid subfamily of Formicidae: Moreau, Bell, et al. 2006: 102.
- Dorylinae as formicoid dorylomorph subfamily of Formicidae: Brady, Schultz, et al. 2006: 18173; Ward, 2007CWard, P. S. 2007C. Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae). Zootaxa 1668:549-563. [2007-12-21] PDF 132572: 555.
- Dorylinae as senior synonym of Acanthostichini, Aenictinae, Aenictogitoninae, Cerapachyinae (Eusphinctinae and Lioponerini), Cheliomyrmecini, Cylindromyrmecini, Ecitoninae, Leptanilloidinae: Brady, Fisher, Schultz & Ward, 2014Brady, S. G.; Fisher, B. L.; Schultz, T. R.; Ward, P. S. 2014. The rise of army ants and their relatives: diversification of specialized predatory doryline ants. BMC Evolutionary Biology 2014 14:93, 14 pp. (doi:10.1186/1471-2148-14-93). PDF 142599: 5.
Subfamily, tribe and genus references
[Note: from Bolton 2003; letters after publication dates may not correspond.]Shuckard, 1840: 195 (genera key); Roger, 1863b: 20, 41, 42 (Typhlopone, Anomma, Dorylus, Rhogmus,Dichthadia catalogues); Smith, F. 1858b: 110, 112 (Typhlopone, Anomma diagnoses); Mayr, 1863: 394, 407,408, 453, 457 (Anomma, Dichthadia, Dorylus, Rhogmus, Typhlopone catalogues); Mayr, 1865: 16, 17(Dorylinae, Typhlopone, Anomma, Dorylus, Rhogmus, Dichthadia diagnoses); Mayr, 1867a: 91 (Typhloponediagnosis); Fore!, 1878: 365 (diagnosis); Andre, 1882c: 251 (Europe & Algeria species key); Dalla Torre,1893: 8, 9 (Rhogmus, Anomma, Dory ius) catalogues); Emery, 1895e: 699, 701, 706; 764 (diagosis genus,subgenera, all species key; diagnoses subfamily & tribe); Emery, 1896b: 174 (genera key); Fore!, 1901a: 462(subgenera key); Bingham, 1903: 2 (diagnosis, India, Sri Lanka & Burma species key); Ashmead, 1906: 22,25, 26 (subfamilies, tribes & genera keys); Wheeler, W.M. 1910d: 137 (diagnosis); Emery, 1910b: 4(diagnosis, tribes, key, catalogue); Emery, 1910b: 8 (D. (Dichthadia) diagnosis, catalogue); Emery, 1910b: 9(D. (Dorylus) diagnosis, catalogue); Emery, 1910b: 10 (D. (Anomma) diagnosis, catalogue); Emery, 1910b:12 (D. (Typhlopone) diagnosis, catalogue); Emery, 1910b: 13 (D. (Rhogmus) diagnosis, catalogue); Emery,1910b: 14 (D. (Alaopone) diagnosis, catalogue); Santschi, 1912: 154 (D. (Anomma) species key); Arnold,1915: 113, 114, 115 (tribes key, subgenera key, South Africa D. (Dorylus) species key); Fore!, 1917: 239(synoptic classification); Santschi, 1919a: 231 (D. (Rhogmus) males key); Fore!, 1921: 135 (diagnosis);Wheeler, W.M. 1922a: 39, 41, 633, 727 (subfamily & genus diagnoses, tribes & subgenera key, Afrotropicalcatalogue); Morley, 1939: 114 (phylogeny); Santschi, 1939a: 152 (D. (Alaopone) species, males key); Brown& Nutting, 1950: 123 (venation & phylogeny); Chapman & Capco, 1951: 9 (Asia checklist); Brown, 1954b:28 (phylogeny); Raignier & Boven, 1955: 86, 114 (D. (Anomma) species key); Borgmeier, 1955: 57 (tribeskey); Eisner, 1957: 478 (proventriculus morphology); Wilson, 1964: 436 (Indo-Australian species key);Gotwald, 1969: 49 (mouthparts morphology); Wheeler, G.C. & Wheeler, J. 1972: 36 (diagnosis); Kempf,1972a: 265 (Neotropical, synoptic classification); Boven, 1972: 142 (D. (Anomma) queens key; Brown,1973b: 166 (genera & distribution); Boven, 1975: 196 (D. (Dorylus) queens key; Wheeler, G.C. & Wheeler,J. 1976: 46 (larvae, review & synthesis); Smith, D.R. 1979: 1326 (North America catalogue); Gotwald &Burdette, 1981: 78 (phylogeny); Gotwald, 1982: 168 (Dorylus subgenera key); Snelling, 1981: 390 (synoptic classification); Wheeler, G.C. & Wheeler, J. 1985: 256 (synoptic classification); Billen, 1986: 168 (Dufour'sgland); Dlussky & Fedoseeva, 1988: 79 (synoptic classification); Holldobler & Wilson, 1990: 9 onward(synoptic classification, world genera, keys); Bolton, 1990c: 1357 (diagnosis, morphology, phylogeny);Baroni Urbani, Bolton & Ward, 1992: 316 (phylogeny); Xu, 1994a: 118 (China species key); Bolton, 1994:35 (diagnosis, synoptic classification); Bolton, 1995a: 1049 (census); Bolton, 1995b: 177 (catalogue);Holldobler, Obermayer & Peeters, 1996: 158 (metatibial gland); Brady, Fisher, Schultz & Ward, 2014Brady, S. G.; Fisher, B. L.; Schultz, T. R.; Ward, P. S. 2014. The rise of army ants and their relatives: diversification of specialized predatory doryline ants. BMC Evolutionary Biology 2014 14:93, 14 pp. (doi:10.1186/1471-2148-14-93). PDF 142599 (phylogeny); Boudinot, 2015Boudinot, B. E. 2015. Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): a new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages. European Journal of Taxonomy 120:1-62. PDF 142740: 48 (male diagnosis); Fisher & Bolton, 2016Fisher, B. L.; Bolton, B. 2016. Ants of Africa and Madagascar A Guide to the Genera. Berkeley: University of California Press, i-ix, 1-503. [2016-08-00] 142963: 44 (worker diagnosis); Borowiec, 2016Borowiec, M. L. 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608:1-280. [2016-08-04] 10.3897/zookeys.608.9427 PDF 142961: 1 (subfamily revision, worker and male keys)
†fossil
valid
synonym
subspecies
unidentifiable/excluded from Formicidae
unavailable/nomen nudum
homonym
original combination