Aphaenogaster rugulosa valid Japan. Palearctic. Watanabe, H.; Yamane, S. 1999. New species and new status in the genus Aphaenogaster (Formicidae) from Japan. Pp. 728-736 in: Yamane, S.; Ikudome, S.; Terayama, M. 1999. Identification guide to the Aculeata of the Nansei Islands, Japan. Sapporo: Hokkaido University Press, xii + 831 pp. 132581