<ref name="Ali_et_al_1987">{{cite journal |first1=M. F. |last1=Ali |first2=E. D. |last2=Morgan |first3=A. B. |last3=Attygalle |first4=J. P. J. |last4=Billen |year=1987 |title=Comparison of Dufour gland secretions of two species of ''Lepthothorax'' ants (Hymenoptera: Formicidae) |url= |journal=Zeitschrift für Naturforschung. C, Journal of Biosciences |publisher= |volume=42 |issue= |pages=955–960 |doi= }}</ref>