<ref name="Arnol'di_1977">{{cite journal |first1=K. V. |last1=Arnol'di |year=1977 |title=Review of the harvester ants of the genus ''Messor'' (Hymenoptera, Formicidae) in the fauna of the USSR. [In Russian.] |url=https://antcat.org/documents/112/6941.pdf |journal=Zoologicheskii Zhurnal |publisher= |volume=56 |issue= |pages=1637–1648 |doi= }}</ref>