<ref name="Boomsma_et_al_1987">{{cite journal |first1=J. J. |last1=Boomsma |first2=A. A. |last2=Mabelis |first3=M. G. M. |last3=Verbeek |first4=E. C. |last4=Los |year=1987 |title=Insular biogeography and distribution ecology of ants on the Frisian islands |url= |journal=Journal of Biogeography |publisher= |volume=14 |issue= |pages=21–37 |doi= }}</ref>