<ref name="Brian_&_Brian_1955">{{cite journal |first1=M. V. |last1=Brian |first2=A. D. |last2=Brian |year=1955 |title=On the two forms ''macrogyna'' and ''microgyna'' of the ant ''Myrmica rubra'' L |url= |journal=Evolution |publisher= |volume=9 |issue= |pages=280–290 |doi= }}</ref>