<ref name="Brown_&_Wilson_1959">{{cite journal |first1=W. L., Jr. |last1=Brown |first2=E. O. |last2=Wilson |year=1959 |title=The evolution of the dacetine ants |url=https://antcat.org/documents/499/6781.pdf |journal=Quarterly Review of Biology |publisher= |volume=34 |issue= |pages=278–294 |doi= }}</ref>