<ref name="Collingwood_&_Hughes_1987">{{cite journal |first1=C. A. |last1=Collingwood |first2=J. |last2=Hughes |year=1987 |title=Ant species in Yorkshire, England |url=https://antcat.org/documents/744/12884.pdf |journal=Naturalist (Leeds) |publisher= |volume=112 |issue= |pages=95–101 |doi= }}</ref>