<ref name="Dlussky_1965">{{cite journal |first1=G. M. |last1=Dlussky |year=1965 |title=Entomological research in Kirghizia. Three new species of ants from Kirghizia. [In Russian.] |url=https://antcat.org/documents/878/5733.pdf |journal=Entomologicheskie Issledovaniya v Kirgizii |publisher= |volume=4 |issue= |pages=27–33 |doi= }}</ref>