<ref name="Donisthorpe_1909">{{cite journal |first1=H. |last1=Donisthorpe |year=1909 |title=''Formica sanguinea'', Latr. at Bewdley, with an account of a slave-raid, and description of two gynandromorphs |url= |journal=Zoologist |publisher= |volume=1909 |issue= |pages=463–466 |doi= }}</ref>