<ref name="Ford_&_Forbes_1984">{{cite journal |first1=F. C. |last1=Ford |first2=J. |last2=Forbes |year=1984 |title=Histology of the male reproductive systems in the adults and pupae of two Doryline ants, ''Dorylus'' (''Anomma'') ''wilverthi'' Emery and ''D.'' (''A''.) ''nigricans'' Illiger (Hymenoptera: Formicidae) |url= |journal=Journal of the New York Entomological Society |publisher= |volume=91 |issue= |pages=355–376 |doi= }}</ref>