<ref name="Jessen_et_al_1979">{{cite journal |first1=K. |last1=Jessen |first2=U. |last2=Maschwitz |first3=M. |last3=Hahn |year=1979 |title=Neue Abdominaldrüsen bei Ameisen. I. Ponerini (Formicidae: Ponerinae) |url= |journal=Zoomorphologie |publisher= |volume=94 |issue= |pages=49–66 |doi= }}</ref>