<ref name="Kôno_&_Sugihara_1939">{{cite journal |first1=H. |last1=Kôno |first2=Y. |last2=Sugihara |year=1939 |title=The ants of fir and pine forests in Japan. [In Japanese.] |url=https://antcat.org/documents/5030/4633.pdf |journal=Transactions of the Kansai Entomological Society |publisher= |volume=8 |issue= |pages=8–14 |doi= }}</ref>