<ref name="Marsh_1986">{{cite journal |first1=A. C. |last1=Marsh |year=1986 |title=Checklist, biological notes and distribution of ants in the central Namib Desert |url=https://antcat.org/documents/2110/4430.pdf |journal=Madoqua |publisher= |volume=14 |issue= |pages=333–344 |doi= }}</ref>