<ref name="Peeters_&_Crewe_1985">{{cite journal |first1=C. |last1=Peeters |first2=R. M. |last2=Crewe |year=1985 |title=Worker reproduction in the ponerine ant ''Ophthalmopone berthoudi'': an alternative form of eusocial organization |url=https://antcat.org/documents/2327/4235.pdf |journal=Behavioral Ecology and Sociobiology |publisher= |volume=18 |issue= |pages=29–37 |doi= }}</ref>