<ref name="Snyder_1992">{{cite journal |first1=L. E. |last1=Snyder |year=1992 |title=The genetics of social behavior in a polygynous ant |url=https://antcat.org/documents/8437/snyder_1992_naturwissenschaften_genetics_polygynous_ant.pdf |journal=Naturwissenschaften |publisher= |volume=79 |issue= |pages=525–527 |doi= }}</ref>