<ref name="Sugihara_1933">{{cite journal |first1=Y. |last1=Sugihara |year=1933 |title=Hymenoptera-Fauna in Province Tosa. Formicidae. [In Japanese and English.] |url= |journal=Kansai Konchu Zasshi |publisher= |volume=1 |issue= |pages=79–86 |doi= }}</ref>