<ref name="Villet_1991">{{cite journal |first1=M. H. |last1=Villet |year=1991 |title=Reproduction and division of labour in ''Platythyrea'' cf. ''cribrinodis'' (Gerstaecker 1858) (Hymenoptera Formicidae): comparisons of individuals, colonies and species |url=https://antcat.org/documents/8352/villet-1991-reproduction-and-division-of-labour.pdf |journal=Tropical Zoology |publisher= |volume=4 |issue= |pages=209–231 |doi= }}</ref>