<ref name="Wheeler_&_Hölldobler_1986">{{cite journal |first1=D. E. |last1=Wheeler |first2=B. |last2=Hölldobler |year=1986 |title=Cryptic phargmosis: the structural modifications |url= |journal=Psyche (Cambridge) |publisher= |volume=92 |issue= |pages=337–353 |doi= }}</ref>