<ref name="Wheeler_&_Wheeler_1973">{{cite journal |first1=G. C. |last1=Wheeler |first2=J. |last2=Wheeler |year=1973 |title=Ant larvae of four tribes: second supplement (Hymenoptera: Formicidae: Myrmicinae) |url=https://antcat.org/documents/3333/3115.pdf |journal=Psyche (Cambridge) |publisher= |volume=80 |issue= |pages=70–82 |doi= }}</ref>