<ref name="Wheeler_&_Wheeler_1986">{{cite journal |first1=G. C. |last1=Wheeler |first2=J. |last2=Wheeler |year=1986 |title=The larva of ''Proatta'' (Hymenoptera: Formicidae) |url=https://antcat.org/documents/3355/3218.pdf |journal=Psyche (Cambridge) |publisher= |volume=92 |issue= |pages=447–450 |doi= }}</ref>