<ref name="Wheeler_&_Wheeler_1986">{{cite journal |first1=G. C. |last1=Wheeler |first2=J. |last2=Wheeler |year=1986 |title=Supplementary studies on ant larvae: Myrmicinae (Hymenoptera: Formicidae) |url=https://antcat.org/documents/3359/3222.pdf |journal=Journal of the New York Entomological Society |publisher= |volume=94 |issue= |pages=489–499 |doi= }}</ref>