<ref name="Wheeler_1906">{{cite journal |first1=W. M. |last1=Wheeler |year=1906 |title=An ethological study of certain maladjustments in the relations of ants to plants |url=https://antcat.org/documents/3425/10538.pdf |journal=Bulletin of the American Museum of Natural History |publisher= |volume=22 |issue= |pages=403–418 |doi= }}</ref>