<ref name="Wheeler_1931">{{cite journal |first1=W. M. |last1=Wheeler |year=1931 |title=Concerning some ant gynandromorphs |url=http://hymfiles.biosci.ohio-state.edu/pdfs-osuc/9555/9555.pdf |journal=Psyche (Cambridge) |publisher= |volume=38 |issue= |pages=80–85 |doi= }}</ref>