<ref name="Wheeler_1934">{{cite journal |first1=W. M. |last1=Wheeler |year=1934 |title=An Australian ant of the genus ''Leptothorax'' Mayr |url=https://antcat.org/documents/3623/3434.pdf |journal=Psyche (Cambridge) |publisher= |volume=41 |issue= |pages=60–62 |doi= }}</ref>