<ref name="Wilson_1958">{{cite journal |first1=E. O. |last1=Wilson |year=1958 |title=Studies on the ant fauna of Melanesia III. ''Rhytidoponera'' in western Melanesia and the Moluccas. IV. The tribe Ponerini |url=https://antcat.org/documents/3682/3482.pdf |journal=Bulletin of the Museum of Comparative Zoology |publisher= |volume=119 |issue= |pages=303–371 |doi= }}</ref>