<ref name="Yamane_et_al_1985">{{cite journal |first1=S. |last1=Yamane |first2=Y. |last2=Harada |first3=M. |last3=Yano |year=1985 |title=Ant fauna of Tanega-shima Island, the northern Ryukyus (Hymenoptera, Formicidae) |url= |journal=Memoirs of the Kagoshima University Research Center for the South Pacific |publisher= |volume=6 |issue= |pages=166–173 |doi= }}</ref>