<ref name="Yasumatsu_1935">{{cite journal |first1=K. |last1=Yasumatsu |year=1935 |title=Further notes on the hymenopterous fauna of the Yaeyama Group |url=http://hymfiles.biosci.ohio-state.edu/pdfs-osuc/23257/23257.pdf |journal=Annotationes Zoologicae Japonenses |publisher= |volume=15 |issue= |pages=33–45 |doi= }}</ref>