<ref name="Yasuno_1963">{{cite journal |first1=M. |last1=Yasuno |year=1963 |title=The study of the ant population in the grassland at Mt. Hakkôda. I. The distribution and nest abundance of ants in the grassland |url= |journal=Ecological Review |publisher= |volume=16 |issue= |pages=83–91 |doi= }}</ref>