<ref name="Young_1986">{{cite journal |first1=A. M. |last1=Young |year=1986 |title=Notes on the distribution and abundance of ground- and arboreal-nesting ants (Hymenoptera: Formicidae) in some Costa Rican cacao habitats |url= |journal=Proceedings of the Entomological Society of Washington |publisher= |volume=88 |issue= |pages=550–571 |doi= }}</ref>