<ref name="Borowiec_et_al_2009">{{cite journal |first1=M. L. |last1=Borowiec |first2=L. |last2=Borowiec |first3=S. |last3=Csősz |first4=A. |last4=Radchenko |year=2009 |title=Ants collected during 2006 Polish expedition to Kyrgyzstan (Hymenoptera: Formicidae) |url=https://antcat.org/documents/4266/Borowiec_et_al_2009_Genus ants of Kyrgyzstan.pdf |journal=Genus (Wroclaw) |publisher= |volume=20 |issue= |pages=367–379 |doi= }}</ref>