<ref name="Brady_et_al_2006">{{cite journal |first1=S. G. |last1=Brady |first2=T. R. |last2=Schultz |first3=B. L. |last3=Fisher |first4=P. S. |last4=Ward |year=2006 |title=Evaluating alternative hypotheses for the early evolution and diversification of ants |url=https://antcat.org/documents/3814/21124.pdf |journal=Proceedings of the National Academy of Sciences of the United States of America |publisher= |volume=103 |issue= |pages=18172–18177 |doi= }}</ref>