<ref name="Csősz_2003">{{cite journal |first1=S. |last1=Csősz |year=2003 |title=A key to the Ponerinae species of the Carpathian Basin (Hymenoptera: Formicidae) |url=https://antcat.org/documents/8527/annals_hnhm_2003_vol_95_147.pdf |journal=Annales Historico-Naturales Musei Nationalis Hungarici |publisher= |volume=95 |issue= |pages=147–160 |doi= }}</ref>