<ref name="Gobin_et_al_2003">{{cite journal |first1=B. |last1=Gobin |first2=F. |last2=Ito |first3=J. P. J. |last3=Billen |year=2003 |title=The subepithelial gland in ants: a novel exocrine gland closely associated with the cuticle surface |url=https://antcat.org/documents/4396/Gobin_et_al_2003_Acta_Zool.pdf |journal=Acta Zoologica (Copenhagen) |publisher= |volume=84 |issue= |pages=285–291 |doi= }}</ref>