<ref name="Gronenberg_&_Hölldobler_1999">{{cite journal |first1=W. |last1=Gronenberg |first2=B. |last2=Hölldobler |year=1999 |title=Morphologic representation of visual and antennal information in the ant brain |url=https://antcat.org/documents/4415/Gronenberg&Hölldobler_1999_J_Comp_Neurol.pdf |journal=Journal of Comparative Neurology |publisher= |volume=412 |issue= |pages=229–240 |doi= }}</ref>