<ref name="Lorite_et_al_2002">{{cite journal |first1=P. |last1=Lorite |first2=J. A. |last2=Carrillo |first3=A. |last3=Tinaut |first4=T. |last4=Palomeque |year=2002 |title=Comparative study of satellite DNA in ants of the ''Messor'' genus |url=https://antcat.org/documents/4514/Lorite_et_al_2002_Gene.pdf |journal=Gene (Amsterdam) |publisher= |volume=297 |issue= |pages=113–122 |doi= }}</ref>