<ref name="Mackay_2000">{{cite journal |first1=W. P. |last1=Mackay |year=2000 |title=A review of the New World ants of the subgenus ''Myrafant'', (genus ''Leptothorax'') (Hymenoptera: Formicidae) |url=https://antcat.org/documents/3954/14659.pdf |journal=Sociobiology |publisher= |volume=36 |issue= |pages=265–444 |doi= }}</ref>