<ref name="Naumann_et_al_1999">{{cite journal |first1=K. |last1=Naumann |first2=W. B. |last2=Preston |first3=G. L. |last3=Ayre |year=1999 |title=An annotated checklist of the ants (Hymenoptera: Formicidae) of British Columbia |url= |journal=Journal of the Entomological Society of British Columbia |publisher= |volume=96 |issue= |pages=29–68 |doi= }}</ref>