<ref name="Pusch_et_al_2006">{{cite journal |first1=K. |last1=Pusch |first2=J. |last2=Heinze |first3=S. |last3=Foitzik |year=2006 |title=The influence of hybridization on colony structure in the ant species ''Temnothorax nylanderi'' and ''T. crassispinus'' |url=https://antcat.org/documents/4611/Pusch_et_al_2006_Insectes_Soc.pdf |journal=Insectes Sociaux |publisher= |volume=53 |issue= |pages=439–445 |doi= }}</ref>