<ref name="Rigato_&_Scupola_2008">{{cite journal |first1=F. |last1=Rigato |first2=A. |last2=Scupola |year=2008 |title=Two new species of the ''Pyramica gundlachi''-group from Ecuador (Hymenoptera Formicidae) |url=https://antcat.org/documents/4010/21831.pdf |journal=Memoirs on Biodiversity |publisher= |volume=1 |issue= |pages=477–481 |doi= }}</ref>