<ref name="Thorne_&_Traniello_2003">{{cite journal |first1=B. L. |last1=Thorne |first2=J. F. A. |last2=Traniello |year=2003 |title=Comparative social biology of basal taxa of ants and termites |url=https://antcat.org/documents/4723/Thorne&Traniello_2003_ARE.pdf |journal=Annual Review of Entomology |publisher= |volume=48 |issue= |pages=283–306 |doi= }}</ref>