<ref name="Umphrey_&_Danzmann_1998">{{cite journal |first1=G. J. |last1=Umphrey |first2=R. G. |last2=Danzmann |year=1998 |title=Electrophoretic evidence for hybridization in the ant genus ''Acanthomyops'' (Hymenoptera: Formicidae) |url= |journal=Biochemical Systematics and Ecology |publisher= |volume=26 |issue= |pages=431–440 |doi= }}</ref>