<ref name="Wilson_&_Hölldobler_2005">{{cite journal |first1=E. O. |last1=Wilson |first2=B. |last2=Hölldobler |year=2005 |title=The rise of the ants: a phylogenetic and ecological explanation |url=https://antcat.org/documents/4775/Wilson&Hölldobler_2005A_PNAS.pdf |journal=Proceedings of the National Academy of Sciences of the United States of America |publisher= |volume=102 |issue= |pages=7411–7414 |doi= }}</ref>