<ref name="Sundström_et_al_2005">{{cite journal |first1=L. |last1=Sundström |first2=P. |last2=Seppä |first3=P. |last3=Pamilo |year=2005 |title=Genetic population structure and dispersal patterns in ''Formica'' ants — a review |url=https://antcat.org/documents/4848/Sundström_et_al_2005_Ann_Zool_Fenn Population structure and dispersal in Formica - review.pdf |journal=Annales Zoologici Fennici |publisher= |volume=42 |issue= |pages=163–177 |doi= }}</ref>