<ref name="Archibald_et_al_2011">{{cite journal |first1=S. B. |last1=Archibald |first2=K. R. |last2=Johnson |first3=R. W. |last3=Mathewes |first4=D. R. |last4=Greenwood |year=2011 |title=Intercontinental dispersal of giant thermophilic ants across the Arctic during early Eocene hyperthermals |url=https://antcat.org/documents/5398/archibald_et_al_2011_proc_r_soc_b-titanomyrma.pdf |journal=Proceedings of the Royal Society. Series B, Biological Sciences |publisher= |volume=278 |issue= |pages=3679–3686 |doi= }}</ref>