<ref name="Zhou_et_al_2012">{{cite journal |first1=S. |last1=Zhou |first2=Y. |last2=Chen |first3=Z. |last3=Chen |first4=P. |last4=Zhou |first5=D. |last5=Ban |first6=M. |last6=Huang |year=2012 |title=Two new species of the genus ''Leptogenys'' from Guangxi, China (Hymenoptera: Formicidae) |url=https://antcat.org/documents/5942/zhou_et_al_2012_sociobiology_leptogenys_in_china.pdf |journal=Sociobiology |publisher= |volume=59 |issue= |pages=885–892 |doi= }}</ref>