<ref name="Zryanin_2015">{{cite journal |first1=V. A. |last1=Zryanin |year=2015 |title=The first subterranean ant species of the genus ''Meranoplus'' F. Smith, 1853 (Hymenoptera: Formicidae) from Vietnam. Caucasian Entomological Bulletin. |url=https://antcat.org/documents/6640/zryanin__v-a-_2015-_the_first_subterranean_ant_species_of_the_genus_meranoplus_from_vietnam.pdf |journal=Caucasian Entomological Bulletin |publisher= |volume=11 |issue= |pages=153–160 |doi= }}</ref>