<ref name="Werner_et_al_2018">{{cite journal |first1=P. |last1=Werner |first2=P. |last2=Bezděčka |first3=K. |last3=Bezděčková |first4=P. |last4=Pech |year=2018 |title=An updated checklist of the ants (Hymenoptera, Formicidae) of the Czech Republic |url=https://antcat.org/documents/8071/werner_et_al_2018_acta_rer_natur_checklist_of_ants_of_czech_republic.pdf |journal=Acta rerum naturalium |publisher= |volume=22 |issue= |pages=5–12 |doi= }}</ref>