<ref name="Zhuang_et_al_2022">{{cite journal |first1=Y. |last1=Zhuang |first2=W. |last2=Xu |first3=G. |last3=Zhang |first4=H. |last4=Mai |first5=X. |last5=Li |first6=H. |last6=He |first7=H. |last7=Ran |first8=Y. |last8=Liu |year=2022 |title=Unparalleled details of soft tissues in a Cretaceous ant |url=https://antcat.org/documents/8403/zhuang_et_al_2022_bmcecolevol_zigrasimecia.pdf |journal=BMC Ecology and Evolution |publisher= |volume=22 |issue= |pages=1–10 |doi=10.1186/s12862-022-02099-2 }}</ref>