<ref name="Peeters_et_al_2017">{{cite journal |first1=C. |last1=Peeters |first2=M. |last2=Molet |first3=C.-C. |last3=Lin |first4=J. |last4=Billen |year=2017 |title=Evolution of cheaper workers in ants: a comparative study of exoskeleton thickness |url=https://antcat.org/documents/8543/peeters_et_al-_bjls_2017.pdf |journal=Biological Journal of the Linnean Society |publisher= |volume=121 |issue= |pages=556–563 |doi=10.1093/biolinnean/blx011 }}</ref>